Thousands of years after a cataclysmic event destroyed civilization, humanity has adapted and evolved a new way of living. Now, gigantic moving cities roam the Earth, ruthlessly devouring small traction towns. Tom Natsworthy (Robert Sheehan), from the lower classes of the great traction city of London, is forced to fight for survival after encountering the dangerous fugitive Hatter Shaw (Hella Hillmer). Two opposites who should not have met, form an unexpected alliance that is destined to change the future.