Ten years after fighting against the invasion of aliens and successfully sealing the wormhole, humans stood up from the ruins. They forgot their fear and indulged in revelry, including Jack (John Boyega), the son of General Steck Pentecost. In the process of stealing abandoned mecha parts, he accidentally encountered Amara (Cailee Spaeny), a girl who assembled a mini mecha on her own. After that, under the arrangement of his sister Asako Mori (Rinko Kikuchi), Jack and Amara received training as mecha pilots at the Magnolia Base in Mo. At the same time, the Shaw Group, headquartered in Shanghai, is committed to replacing traditional mechas with unmanned mechas under the chairmanship of President Shao Liwen (Jing Tian). At this critical moment, a black mecha that is extremely similar to the Revenge Wanderer suddenly appeared and launched a ruthless killing of humans. People found that the distant alien "pioneers" are still ready to move, waiting for the opportunity to destroy the earthlings...